The final result of investigation is: Main cause

The final result of investigation is: Main cause of mailfunction is Q2, but Shortcut on +5 USB voltage to null remained. Then I did something I should do at the first, to look on rear panel USB ports. One of them was slightly demiged with +5V pin sliped to chasis of connector. That was reason of shortcut. I changed whole connector just to be safe.
P.S. Capabilitiy of users to ruin MOBOS are infinite:)